CAMBRIANO

Trilobitas características gerais

 

Sistemática


Redlichiiida


Agnostida


Corynexochiida


Lichida


Phacopida 


Ptychopariida 


Asaphida 


Proetida 

 

Classificação de trilobitas

Ordem Agnostida
Ordem Redlichiida
Ordem Corynexochida
Ordem Lichida
Ordem Phacopida
Ordem Proetida
Ordem Asaphida
Ordem Ptychopariida
Ordem desconhecidas
Texto em inglês temporariamente

Class Trilobita
.....Order Agnostida
..........Suborder Agnostina
...............Superfamily Agnostoidea
....................Family Agnostidae
.........................Subfamily Agnostinae
..............................Agnostus (Agnostus)
...................................A. (A.) pisiformis., Cambrian, Sweden.
..............................Agnostus (Homagnostus)
...................................A. (H.) obesus (U. Cam.)
..............................Acutatagnostus
..............................?Gymnagnostus
..............................Innitagnostus
..............................?Ivshinagnostus
..............................Lotagnostus (Lotagnostus)
...................................L. trisectus (U.Cam.)
..............................Lotagnostus (Eolotagnostus)
..............................Lotagnostus (Distagnostus)
..............................Micragnostus
...................................M. calvus (U.Cam-L.Ord.)
..............................Oncagnostus (Oncagnostus)
...................................O. (O.) grandis (U. Cam.)
..............................?Eurudagnostus = Oncagnostus (Oncagnostus)
..............................Oncagnostus (Kymagnostus)
..............................Oncagnostus (Strictagnostus)
..............................Trilobagnostus
...................................T. innocens (U.Cam.)
.........................Subfamily Ammagnostinae
..............................Ammagnostus (Ammagnostus)
..............................Ammagnostus (Tentagnostus)
..............................Formosagnostus
..............................Hadragnostus
..............................?Kormagnostella
..............................Kormagnostus
...................................K. simplex (U.Cam.)
..............................Proagnostus
.........................Subfamily Glyptagnostinae
..............................Glyptagnostus
...................................Glyptagnostus toreuma (U.Cam.)
..............................Agnostardis
.........................Subfamily Uncertain
..............................Acmarhachis
..............................Acmarhachis typicalis (U.Cam.)
..............................Aistagnostus
..............................Connagnostus
..............................?Idolagnostus
..............................Quadrahomagnostus
....................Family Ptychagnostidae
..............................Ptychagnostus
...................................Ptychagnostus punctuosus (M.Cam.)
..............................Acidusus
..............................Goniagnostus (Goniagnostus)
...................................G. (G.) nathorsti (M.Cam.)
..............................Goniagnostus (Allobodochus)
..............................Goniagnostus (Criotypus)
..............................Lejopyge
...................................Lejopyge laevigata (M.-U.Cam.)
..............................Myrmecomimus
..............................Onymagnostus
..............................Pentagnostus
...................................Pentagnostus anaberensis (M.Cam.)
..............................Tomagnostus
...................................Tomagnostus fissus (M.Cam.)
..............................Triplagnostus (Triplagnostus)
...................................T. (T.) gibbus (M.Cam.)
..............................Triplagnostus (Aotagnostus)
..............................Zeteagnostus
....................Family Spinagnostidae
.........................Subfamily Spinagnostinae
..............................Hypagnostus
...................................Hypagnostus parvifrons (M.Cam.)
..............................Cotalagnostus
...................................Cotalagnostus lens (M.Cam.)
..............................Eoagnostus
...................................Eoagnostus roddyi (L.Cam.)
..............................Euagnostus
...................................Euagnostus opimus (M.Cam.)
..............................Micagnostus
..............................Peratagnostus
..............................Raragnostus
..............................Svenax
..............................Tomagnostella
...................................Tomagnostella exsculpta (M.Camb.)
.........................Subfamily Doryagnostinae
..............................Doryagnostus
...................................Doryagnostus incertus (M.Cam.)
..............................Itagnostus
..............................Rhodotypiscus
.........................Subfamily Uncertain
..............................Pseudoperonopsis
....................Family Peronopsidae
..............................Peronopsis
...................................Peronopsis integra (M.Cam.)
..............................Diplorrhina
..............................Gratagnostus
..............................Peronopsella
....................Family Diplagnostidae
.........................Subfamily Diplagnostinae
..............................Diplagnostus
...................................Diplagnostus planicauda (M.Cam.)
..............................Acadagnostus
...................................Acadagnostus acadicus (M.Cam.)
..............................Baltagnostus
...................................Baltagnostus centerensis (M.-U.Cam.)
..............................Dolichagnostus
..............................?Iniospheniscus
..............................Linguagnostus
...................................Linguagnostus kjerulfi (M.Cam.)
..............................Oedorhachis
...................................Oedorhachis typicalis (U.Cam.)
..............................Tasagnostus
.........................Subfamily Oidalagnostinae
..............................Oidalagnostus
...................................Oidalagnostus trispinifer (M.Cam.)
..............................Cristagnostus
.........................Subfamily Pseudoagnostinae
..............................Pseudagnostus (Pseudagnostus)
...................................P. (P.) cyclopyge (U.Cam.)
..............................Pseudagnostus (Pseudagnostina)
..............................Pseudagnostus (Sulcatagnostus)
...................................P. (S.) securiger (U.Cam.)
..............................Agnostotes (Agnostotes)
..............................Agnostotes (Pseudoglyptagnostus)
..............................Denagnostus
..............................Litagnostus
...................................Litagnostus levisensis (U.Cam.)
..............................Nahannagnostus
..............................Neoagnostus (Neoagnostus)
...................................N. (N.) aspidoides (L.Ord.)
..............................Neoagnostus (Machairagnostus)
...................................N. (M.) tmetus (L.Ord.)
..............................Oxyagnostus
..............................Rhaptagnostus
...................................Rhaptagnostus cyclopygeformis (U.Cam.-L.Ord.)
..............................Trisulcagnostus
..............................Xestagnostus
....................Family Clavagnostidae
.........................Subfamily Clavagnostinae
..............................Clavagnostus
...................................Clavagnostus repandus (M.-U.Cam.)
.........................Subfamily Aspidagnostinae
..............................Aspidagnostus
...................................Aspidagnostus parmatus (U.Cam.)
..............................Triadaspis
.........................Subfamily Uncertain
..............................Utagnostus
....................Family Metagnostidae
..............................Arthrorachis
..............................Anglagnostus
...................................Anglagnostus dux (U.Cam-L.Ord.)
..............................Chatkalagnostus
..............................Corrugatagnostus
...................................Corrugatagnostus perrugatus (L.-M.Ord.)
..............................Dividuagnostus
..............................Galbagnostus
..............................Geragnostella
...................................Geragnostella tullbergi (L.Ord.)
..............................Geragnostus
...................................Geragnostus sidenbladhi (L.Ord.)
..............................Granduloagnostus
..............................Homagnostoides
...................................Homagnostoides ferralsensis (L.Ord.)
..............................Trinodus
...................................Trinodus agnostiformis (Ord.)
....................Family Uncertain
..............................Agnostogonus
..............................Archaeagnostus
...................................Archaeagnostus primageneus (L.Cam.)
..............................Armagnostus
...................................Armagnostus megalaxis (M.Cam.)
..............................Blystagnostus
..............................Ciceragnostus
...................................Ciceragnostus barlowi (M.Cam.)
..............................Delagnostus
..............................Gallagnostus
...................................Gallagnostus geminus (U.Cam-L.Ord.)
..............................Glaberagnostus
..............................Grandagnostus
...................................Grandagnostus vermontensis (M.Cam.)
..............................Hastagnostus
...................................Hastagnostus angustus (M.Cam.)
..............................Leiagnostus
...................................Leiagnostus erraticus (L.Ord.)
..............................Monaxagnostus
..............................Phalagnostus
...................................Phalagnostus nudus (M.Cam.)
..............................Phaldagnostus
..............................Phoidagnostus
..............................Plurinodus
..............................Quadragnostus
...................................Quadragnostus solus (M.Cam.)
..............................Skryjagnostus
..............................Toragnostus
..............................Valenagnostus
...............Superfamily Uncertain
....................Family Phalacromidae
..............................Phalacroma
...................................Phalacroma bibullatum (M.Cam.)
..............................?Dignagnostus
..............................Lisogoragnostus
....................Family Sphaeragnostidae
..............................Sphaeragnostus
...................................Sphaeragnostus similaris (Ord.)
...............Superfamily Condylopygoidea
....................Family Condylopygidae
..............................Condylopyge
...................................Condylopyge rex (M.Cam.)
..............................Pleuroctenium
...................................Pleuroctenium granulatum (M.Cam.)

Nomina nuda

Cenagnostus
Clavagnostotes
Kitatagnostus
Munagnostus
Parapseudagnostus
Pseudagnostella

Rejected Genera:

Discagnostus (in Eodiscida)
Gallagnostoides
Mallagnostus (in Eodiscida)
Pseudotrinodus

..........Suborder Eodiscina
...............Superfamily Eodiscoidea
....................Family Tsunyidiscidae
..............................Tsunyidiscus
....................Family Hebediscidae
..............................Hebediscus
...................................Hebediscus attleborensis (L.Cam.)
..............................Delgadella
...................................Delgadella lusitanica (L.Cam.)
..............................Dicerodiscus
..............................Luvsanodiscus
..............................Natalina
..............................Neopagetina
..............................Parapagetia
..............................Tchernyshevioides
....................Family Calodiscidae
..............................Calodiscus
...................................Calodiscus lobatus (L.Cam.)
..............................Chelediscus
..............................Korobovia
..............................Neocobboldia
...................................Neocobboldia dentata (L.Cam.)
..............................Pseudocobboldia
..............................Sinodiscus
....................Family Weymouthiidae
..............................Weymouthia
...................................Weymouthia nobilis (L.Cam.)
..............................Abakolia
..............................Acidiscus
..............................Acimetopus
..............................Analox
..............................Bathydiscus
..............................Bolboparia
..............................Cephalopyge
..............................Cobboldites
..............................Jinghediscus
..............................Leptochilodiscus
..............................Litometopus
..............................Mallagnostus
..............................Meniscuchus
..............................Ninadiscus
..............................Oodiscus
..............................Runcinodiscus
..............................Semadiscus
..............................Serrodiscus
...................................Serrodiscus serratus (L.Cam.)
...................................Serrodiscus speciosus (L.Cam.)
..............................Stigmadiscus
..............................Tanudiscus
....................Family Yukoniidae
..............................Yukonia
..............................Alaskadiscus
..............................Egyngolia
..............................Ekwipagetia
..............................Hebediscina
..............................Lenadiscus
..............................Yukonides
....................Family Eodiscidae
..............................Eodiscus
...................................Eodiscus punctatus (M.Cam.)
..............................Dawsonia
...................................Dawsonia dawsoni (M.Cam.)
..............................Helepagetia
..............................Kiskinella
..............................Macannaia
..............................Opsidiscus
...................................Opsidiscus bilobatus (M.Cam.)
..............................Pagetia
...................................Pagetia bootes (L.-M.Cam.)
..............................Pagetides
...................................Pagetides elegans (L.Cam.)
...................................Pagetides rupestris (L.Cam.)
..............................Sinopagetia

Genera previously assigned to Eodiscoidea but here considered doubtful or excluded:

Dipharus
Discagnostus
Glabrella
Menodiscus
Miraculaspis
Shivelicus
Triangulaspis

.....Order Redlichiida
..........Suborder Olenellina
...............Superfamily Olenelloidea
....................Family Olenellidae
.........................Subfamily Olenellinae
..............................Olenellus (Olenellus)
..............................Olenellus (Angustolenellus)
..............................Olenellus (Mesolenellus)
..............................Olenellus (Mesonacis)
..............................Olenellus (Paedeumias)
...................................O. (P.) transitans (L.Cam.)
..............................Fremontella
...................................Fremontella halli (L.Cam.)
..............................Mummaspis
.........................Subfamily Biceratopsinae
..............................Biceratops
..............................Peachella
...................................Peachella idingsi (L.Cam.)
.........................Subfamily Bristoliinae
..............................Bristolia
...................................Bristolia bristolensis (L.Cam.)
..............................Arcuolenellus
..............................Bolbolenellus
..............................Nephrolenellus
.........................Subfamily Gabriellinae
..............................Gabriellus
.........................Subfamily Laudoniinae
..............................Laudonia
...................................Laudonia bispinata (L.Cam.)
..............................?Olenelloides
.........................Subfamily Wanneriinae
..............................Wanneria
....................Family Holmiidae
.........................Subfamily Holmiinae
..............................Holmia
..............................Andalusiana
..............................Cambropallas
..............................Elliptocephala
..............................Esmeraldina
..............................Holmiella
..............................Iyouella
..............................Palmettaspis
..............................Schmidtiellus
.........................Subfamily Callaviinae
..............................Callavia
...................................Callavia broeggeri (L.Cam.)
..............................Kjerulfia
...................................Kjerulfia lata (L.Cam.)
...............Superfamily Fallotaspidoidea
....................Family Fallotaspididae
.........................Subfamily Fallotaspidinae
..............................Fallotaspis
..............................Eofallotaspis
..............................Lenallina
..............................Parafallotaspis
..............................Pelmanaspis
..............................Profallotaspis
.........................Subfamily Daguinaspidinae
..............................Daguinaspis
..............................Choubertella
..............................?Wolynaspis
....................Family Archaeaspididae
..............................Archaeaspis
..............................Bradyfallotaspis
..............................Fallotaspidella
..............................Geraldinella
..............................Selindella
....................Family Judomiidae
..............................Judomia
...................................Judomia dzevanovskii (L.Cam.)
..............................Judomiella
..............................Paranevadella
..............................Sinskia
....................Family Neltneriidae
..............................Neltneria
..............................Bondella
....................Family Nevadiidae
..............................Nevadia
..............................Buenellus
..............................Cirquella
..............................Nevadella
..............................Pseudojudomia
...............Superfamily Uncertain
..............................Poletavella
..............................Postfallotaspis

..........Suborder Redlichiina
...............Superfamily Emuelloidea
....................Family Emuellidae
..............................Emuella
..............................Balcoracania
...............Superfamily Redlichioidea
....................Family Redlichiidae
.........................Subfamily Redlichiinae
..............................Redlichia (Redlichia)
..............................Redlichia (Conoredlichia)
..............................Redlichia (Latiredlichia)
..............................Redlichia (Pteroredlichia)
..............................Syndianella
.........................Subfamily Metaredlichiinae
..............................Metaredlichia
..............................Bornemannaspis
..............................Breviredlichia
..............................Iglesiella
..............................Jingyangia
..............................Maopingaspis
..............................?Nebidella
..............................Parazhenbaspis
..............................Pseudoredlichia
..............................Sardoredlichia
..............................Ushbaspis
..............................Xela
..............................Zhenbaspis
.........................Subfamily Neoredlichiinae
..............................Neoredlichia
..............................Leptoredlichia
..............................L. (Leptoredlichia)
..............................L. (Xenoredlichia)
..............................?Olgaspis
.........................Subfamily Pararedlichiinae
..............................Eoredlichia
..............................E. (Eoredlichia)
..............................E. (Pachyredlichia)
..............................?Irgitkhemia
..............................Lemdadella
..............................Ningqiangaspis
..............................?Redlichiops
.........................Subfamily Wutingaspinae
..............................Wutingaspis
..............................Chaoaspis
..............................Chengjiangaspis
..............................Kepingaspis
..............................Kuanyangia
..............................K. (Kuanyangia)
..............................K. (Sapushania)
..............................Sardaspis
..............................?Wengangaspis
..............................?Yorkella
....................Family Dolerolenidae
.........................Subfamily Doleroleninae
..............................Dolerolenus
..............................D. (Dolerolenus)
..............................D. (Malungia)
.........................?Subfamily Paramalungiinae
..............................Paramalungia
....................Family Yinitidae
..............................Yinites
..............................Drepanopyge
..............................Drepanuroides
..............................Meitanella
..............................Paokannia
..............................Parapaokannia
..............................Parayinites
..............................Pseudopaokannia
..............................Quingkouia
..............................Yunnanaspidella
..............................Yunnanaspis
....................Family Mayiellidae
..............................Mayiella
..............................Qiaodiella
..............................Qiaotingaspis
....................Family Gigantopygidae
.........................Subfamily Gigantopyginae
..............................Gigantopygus
.........................Subfamily Yiliangellinae
..............................Yiliangella
..............................Yiliangellina
..............................Zhangshania
....................Family Saukiandidae
.........................Subfamily Saukiandinae
..............................Saukianda
..............................S. (Saukianda)
..............................S. (Pseudosaukianda)
..............................Longianda
.........................Subfamily Despujolsiinae
..............................Despujolsia
.........................Subfamily Resseropinae
..............................Perrector
..............................P. (Perrector)
..............................P. (Resserops)
..............................P. (Richterops)
..............................Australaspis
..............................Clariondia
..............................Dolerolichia
..............................Eops
..............................Pareops
..............................Realaspis
....................Family Metadoxididae
..............................Metadoxides
..............................?Churkinia
..............................Enantiaspis
..............................Fuminaspis
..............................Hongshiyanaspis
..............................Minusinella
..............................?Onaraspis
....................Family Abadiellidae
..............................Abadiella
..............................Guangyuanaspis
..............................Lunolenus
..............................Malongocephalus
..............................Parabadiella
..............................Shaanxia
..............................Sibiriaspis
....................Family Kueichowiidae
..............................Kueichowia
..............................Shatania
....................Family Menneraspidae
..............................Menneraspis
....................Family Redlichinidae
..............................Redlichina
..............................Asthenaspis
..............................Kolbaspis
..............................Parasayanaspis
..............................Sajanaspis
..............................Tungusella
....................Family Chengkouaspidae
..............................Chengkouaspis
..............................Belliceps
..............................Bulaiaspis
..............................Inella
..............................Pseudoresserops
..............................Terechtaspis
....................Family Uncertain
..............................Akbashichia
..............................Elegestina
..............................Elganellus
..............................Iolgia
..............................Paratungusella
..............................Sarassina

Unrecognizable redlichioid genera:

Eomalungia
Fandianaspis
Guangyuania
Longmenshania
Micangshania
Parawutingaspis
Pseudowutingaspis
Runnania
Shangsiaspis
Xingzishania
Yiliangella (Pseudoyiliangella)

...............Superfamily Paradoxidoidea
....................Family Paradoxididae
..............................Paradoxides
..............................Acadoparadoxides
..............................A. (Acadoparadoxides)
..............................A. (Baltoparadoxides)
..............................Anabaraceps
..............................Anabaraspis
..............................Eccaparadoxides
..............................Hydrocephalus
..............................Plutonides
..............................?Primoriella
..............................Schagonaria
....................Family Centropleuridae
..............................Centropleura
..............................C. (Centropleura)
..............................C. (Beishanella)
..............................Anopolenus
..............................Clarella
..............................Luhops

.....Order Corynexochida
..........Suborder Corynexochina
...............Superfamily Corynexochoidea
....................Family Corynexochidae
....................Family Cheiruroididae
....................Family Chenghuiidae
....................Family Dorypygidae
....................Family Ogygopsidae
....................Family Oryctocephalidae
....................Family Dolichometopidae
....................Family Edelsteinaspidae
....................Family Jakutidae
....................Family Longduiidae
....................Family Zacanthoididae
....................Family Dinesidae

..........Suborder Illaenina
...............Superfamily Illaenoidea
....................Family Styginidae (Scutelluidae)
..............................Breviscutellum
..............................Paralejurus
....................Family Phillipsinellidae
....................Family Illaenidae
....................Family Tsinaniidae

..........Suborder Leiostegiina
...............Superfamily Leiostegioidea
....................Family Leiostegiidae
....................Family Pagodiidae
....................Family Kaolishaniidae
....................Family Cheilocephalidae
....................Family Lecanopygidae (Illaenuridae) (?)
....................Family Shirakellidae (?)

.....Order Lichida
..........Suborder Lichina
...............Superfamily Lichoidea
....................Family Lichidae
..............................Arctinurus
....................Family Lichakelaphidae
...............Superfamily Odontopleurida
....................Family Odontopleuridae
..............................Kettneraspis
....................Family Selenopeltidae
..............................Dicranurus
...............Superfamily Dameselloidea
....................Family Damesellidae

.....Order Phacopida
..........Suborder Calymenina
...............Superfamily Calymenoidea
....................Family Calymenidae
..............................Calymene
..............................Diacalymene
....................Family Homalonotidae
..............................Trimerus (Dipleura)
....................Family Carmonidaedae
....................Family Bathycheilidae

..........Suborder Phacopina
...............Superfamily Phacopoidea
....................Family Phacopidae
..............................Kainops
..............................Phacops
..............................Reedops
..............................Viaphacops
....................Family Pterygometopidae
..............................Calyptaulax
...............Superfamily Dalmanitoidea
.....................Family Dalmanitidae
..............................Dalmanites
..............................Huntonia
.....................Family Prosopiscidae
.....................Family Diaphanometopidae
...............Superfamily Acastoidea
.....................Family Acastidae
..........................Subfamily Acastavinae
...............................Acastava
...............................Acastella
...............................Acastellina
...............................Acastocephala
...............................Pelitlina
...............................Centauropyge
...............................Mimocryphaeus
...............................Chimaerastella
...............................Acastopyge
..........................Subfamily Acastinae
..........................Subfamily Asteropyginae.

..............................Alcaldops
..............................Armorigreenops
..............................Asteropyge
..............................Bellacartwrightia
..............................Bradocryphaeus
..............................Braunops
..............................Breizops
..............................Coltraneia
..............................Comura
..............................Cryphaeus
..............................Cryphina
..............................Delocare
..............................Deloops
..............................Destombesina
..............................Echinopyge
..............................Greenops
..............................Hallandclarkeops
..............................Heliopyge
..............................Hollardops
..............................Kayserops
..............................Kennacryphaeus
..............................Metacanthina
..............................Modellops
..............................Neocalmonia
..............................Neocalmonia (Hexacosta)
..............................Neocalmonia (Radiopyge)
..............................Neometacanthus
..............................Paracryphaeus
..............................Pelitlina
..............................Philipsmithiana
..............................Philonyx
..............................Pilletina
..............................Proacanthina
..............................Pseudocryphaeus
..............................Psychopyge
..............................Quadratispina
..............................Rhenops
..............................Stummiana
..............................Tolkienia
..............................Treveropyge
.....................Family Calmoniidae

..........Suborder Cheirurina
...............Superfamily Cheiruroidea
....................Family Cheiruridae
..............................Ceraurus
...................................C. pleurexanthemus
....................Family Pliomeridae
....................Family Pilekiidae
....................Family Encrinuridae
....................Family Staurocephalidae
....................Family Hammatocnemidae

.....Order Proetida
..........Suborder Proetina
...............Superfamily Proetoidea
....................Family Proetidae
..............................Basidechenella
..............................Dechenella
....................Family Phillipsiidae
...............Superfamily Aulacopleuroidea
....................Family Aulacopleuridae
....................Family Brachymetopidae
....................Family Rorringtoniidae
...............Superfamily Bathyuroidea
....................Family Bathyuridae
....................Family Dimeropygidae
....................Family Celmidae
....................Family Lecanopygidae (Illaenuridae) (?)
....................Family Glaphuridae
....................Family Holotrachelidae
....................Family Telephinidae

.....Order Asaphida
..........Suborder Asaphina
...............Superfamily Anomocaroidea
....................Family Anomocaridae
....................Family Pterocephaliidae (includes Housiinae)
....................Family Parabolinoididae
....................Family Dikelokephalinidae
...............Superfamily Asaphoidea
....................Family Asaphidae
..............................Homotelus
..............................Pseudogygites
....................Family Ceratopygidae
...............Superfamily Dikelokephaloidea
....................Family Dikelocephalidae
....................Family Saukiidae
....................Family Ptychaspididae
....................Family Eurekiidae
....................Family Loganellidae
...............Superfamily Remopleuridiodea
....................Family Remopleurididae
....................Family Kainellidae
....................Family Opipeuteridae
....................Family Bohemillidae
....................Family Auritamiidae
....................Family Idahoiidae
....................Family Hungaiidae
...............Superfamily Cyclopygoidea
....................Family Cyclopygidae
....................Family Taihungshaniidae
....................Family Nileidae
...............Superfamily Trinucleioidea
....................Family Trinucleidae
....................Family Dionididae (Tongxinaspididae) (?)
....................Family Orometopidae (includes Myindidae, Hapalopleuridae, Jegorovaiidae)
....................Family Raphiophoridae
....................Family Alsataspididae
....................Family Liostracinidae
...............Superfamily Uncertain
....................Family Rhyssometopidae (includes Mapaniidae, Plectriferidae)
....................Family Monkaspididae (Chelidonocephalidae) (?)

 

.....Order Ptychopariida
..........Suborder Ptychopariina
...............Superfamily Ellipsocephaloidea
....................Family Ellipsocephalidae
....................Family Protolenidae...............Superfamily Ptychoparioidea
....................Family Ptychopariidae
....................Family Marjumiidae
....................Family Solenopleuridae
....................Family Conocoryphidae
....................Family Dokimocephalidae
....................Family Onchonotopsidae
....................Family Nepeiidae
....................Family Crepicephalidae
....................Family Lonchocephalidae
....................Family Kingstoniidae
....................Family Shumardiidae
....................Family Asaphiscidae
....................Family Coosellidae
....................Family Elviniidae (includes Komaspididae)
....................Family Cedariidae
....................Family Norwoodiidae
....................Family Menomoniidae
....................Family Bolaspididae
....................Family Papyriaspididae
....................Family Emmrichellidae
....................Family Diceratocephalidae
....................Family Tengfengiidae
....................Family Lisaniidae
....................Family Inouyiidae
....................Family Wuaniidae
....................Family Lorenzellidae
....................Family Proasaphiscidae
....................Family Ignotogregatidae
....................Family Holanshaniidae
....................Family Mapaniidae
....................Family Acrocephalitidae

..........Suborder Olenina
...............Superfamily Olenioidea
....................Family Olenidae
..............................Triarthrus

..........Suborder Harpina
...............Superfamily Harpioidea
....................Family Harpetidae
....................Family Harpididae
....................Family Entomaspididae

..........Suborder Uncertain
....................Family Ithyophoridae

..........Polyphyletic Groups
....................Family Catillicephalidae
....................Family Avoninidae
....................Family Raymondinidae
....................Family Plethopeltidae (near Cedariidae - Norwoodiidae above?)

.....Order Uncertain
....................Family Burlingiidae
....................Family Bathynotidae
....................Family Bestjubellidae
....................Family Naraoiidae

Families in need of placement from 1959 Treatise:
....................Family Isocolidae (Asaphida?)
....................Family Missisquoiidae (Leiostegiina?)
....................Family Loganopeltidae (=Harpididae?)
....................Family Koldinioididae (Ptychopariida?)